| 1988-1993 | Chemistry Studies, University of Athens, Greece |
| 1994-1998 |
Ph.D. at Zentrum für Molekulare Biologie Heidelberg (ZMBH), laboratory of Prof. Stefan Jentsch |
| 2000-2005 |
Postdoctoral studies, Institute of Biochemistry, ETH Zurich, laboratory of Prof. Yves Barral |
| 10/2005 - 09/2013 | Junior group leader at the Biochemie Zentrum Heidelberg (BZH) |